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Smart but wholesome Betty embraces her unique appearance and independent attitude in the cutthroat world of New York fashion.

NSA computer technician Jake Foley is transformed into a secret agent after he is accidentally infected with nanobots, based on nanotechnology that reduce the size of a computer to the molecular level, which give him superhuman powers.

When a mysterious stranger washes up on the beach in Miami, he changes the lives of the three generations of Fuentes women forever.

Tucker Burns, a good journalist with a blemish on his record that makes him unemployable, is hired by The World Chronicle, a paranormal tabloid. Although a skeptic, Tucker agrees to investigate the case of a monster attack for the paper.

The assistants whip themselves into a frenzy over who will get the swag when Christina announces she is going to clean out the magazine's closet.

Betty uses her Queens connections to lure Vincent Bianchi, a famous photographer, to shoot for the magazine. When Bianchi insists that Betty meet him at a posh restaurant, she gets a makeover. But things don't quite turn out as planned. In the mean time, Bradford tries to get his hands on the car Fey Sommers' died in and Wilhelmina primps for a big date.

Betty loses a book, which is the complete new edition of the magazine and is containing un-retouched photos of a popular actress. She has to find the book before the photos get out to the rest of the press. Meanwhile, Wilhelmina plots to use the snafu to bring Daniel down and Bradford Meade secretively inquires about Fey Sommers' final effects.

After Mode is forced to abandon its big Christmas photo shoot because another magazine has copied the idea, Betty forms an uneasy alliance with Marc and Amanda. The three fear they may have leaked the idea to someone at the rival mag. Meanwhile, Daniel and Wilhelmina reluctantly work together to come up with a new photo spread as Mode's deadline looms, and Betty's nephew, Justin, is oblivious to the turmoil as he spends a day at the magazine for a class project.

It's Halloween and Betty is in for a series of frightful experiences, ranging from showing up dressed for the Mode costume contest . . . only to learn there isn't one, to dealing with her growing attraction to Henry, while a jealous Walter fumes -- and, worst of all, tracking down a watch Daniel left at one of his many girlfriends' apartments (he just can't remember which one). Meanwhile, Daniel pays a visit to his mom in rehab, and Wilhelmina is horrified to learn that she can't fit into a designer gown

To his dismay, Daniel must attend his family's Thanksgiving gathering at the Country Club. Meanwhile, Amanda and Marc create their own gathering at Mode and there is also a dinner at Betty's house where Santos, Justin's father, is in attendance. Meanwhile, Wilhelmina also hosts a Thanksgiving dinner as well but her feast doesn't look as good as it does in magazine pictures.

Sofia Reyes spreads herself in Daniel's office to prove her power. But, instead of paying attention to him Sofia is interested in Betty's opinions and ideas. Fearing that his assistant is not challenged enough by her job, Daniel sends Betty off to write a review about a hotel where she is supposed to stay at over the weekend. Actually, Betty has planned to spend the weekend with Walter in Atlantic City, but now has to cancel this trip. When, Walter pays her a surprise visit at the hotel, Betty is at first very relieved to see him. But, when Betty continues her review observations of the hotel, Walter is not very helpful. Meanwhile, at MODE, Daniel, who is still interested in Sofia, continues to try to get her attention, but Sofia seems to be very resistant to his charms. And, Wilhelmina has a meeting with an important client, but has to dress up for this meeting in very unusual clothes for her and as it turns out she has to do much more to please him.. At home, Hilda still tries to get the money together the family needs for a lawyer. Being very desperate, she asks Santos, who gives her the money, but in exchange wants to see his and Hilda's son Justin.

The day after Thanksgiving is no holiday for Betty, who struggles to pull together a last-minute Mode photo shoot with a famous couple and their baby. Meanwhile, Daniel suffers from the after effects of drinking whilst getting over Sofia and ends up spending the holiday with Betty's family and Wilhemina and Marc get 'lost' in Queens!

It's Christmas at Mode as Betty questions her relationship with Walter and her feelings for Henry the accountant. Daniel, on the other hand, is sure of his love for Sofia.

Alexis' shocking announcement causes great commotion at both Mode and the Meade home. Betty tries to maintain the peace and order at Mode, however, she is about to face her own personal turmoil when Henry asks her out on a date and Walter tells her he's got a big surprise in store - he has been offered a promotion in Maryland. Meanwhile, Whilhelmina plots her next move to conquer Mode and Justin gets sent home from school for fighting.

Wilhelmina, with Mark as her right hand, plans who's to stay and who'll be kicked out when she once takes over Daniel's position. Daniel returns to the office after 14 days licking wounds. He immediately rehires Betty as assistant. Hilda has success in delivering her father's cakes and chooses to start a new business without telling him about it before taking in orders.

Sophia Reyes rules Daniel Meade's heart masterfully, getting him to propose to her and winning favor with his parents. Meanwhile as Betty's new boss, Sophia orders her to replace a lead article in her new feminist magazine by one on the social experiences of her, common ugly bird, working in Daniel's fancy fashion magazine, so she goes back there interviewing everybody. Betty finds out that Sophia not only continues to enjoy the services of Hunter, who is actually exotic dancer, but also hired him to play her fiance. Betty deduces Sophia seduced Daniel maliciously, but is too late to tell him before Sophia announces on national TV she only wanted to prove the validity of her book's 60 days plan to hook up any man. She turns his proposal down, which totally destroys his spirit. Meanhile Texan lover Ted LeBeau renders Wilhelmina romantic enough to turn a cold shoulder to her shady boss, until his wife wants him back as father of their children. Betty's dad Ignacio finds he can't get out of the demands of mean Immigration official Constance Grady, who will get him deported even after 30 years of total integration unless he satisfies her.

Betty has her job cut out preparing for Mode's huge New York City's Fashion Week event. Marc and Amanda can't delay getting their hands on something everybody desires, Christina wishes to create her mark in the design world and Daniel calls for Hilda to help her sister. Alexis makes her entrance!

Daniel reconnects with New York City's top lawyer Grace Chin, which he once stood up for a date in college. Betty befriends Charlie, an out-of-towner she meets at the Mode cafeteria but later discovers to be Henry's girlfriend.

Daniel keeps secret from Alexis the fact that he is the one who hired Grace to defend their father, something which Wilhelmina uses to bring the siblings apart. Meanwhile, when invited by Henry to Charlie's birthday party, a desperate Betty asks Dr. Farkas, her orthodontist, to be her date. Finally, Amanda catches the eye of an unusual designer who claims that she is his muse.

Betty is in for some wild drama when Marc's mother, Jean Weiner, stops by Mode, given that the latter doesn't seem to know that her son is gay and Betty acts as his girlfriend to help him keep his secret. Meanwhile, Daniel and Alexis are forced to work together when Claire names them both co-editors-in-chief.

Daniel tells Betty to quit stalking him when she tries to protect him from unauthorized biographer Quincy Combs. Later, Daniel is appalled when he is told his latest bedmate was underage - by her mother.

At a medieval-themed restaurant where Mode assistants celebrate Secretaries' -- er, Administrative Assistants' Day, Henry battles to defend Betty's honor (which leaves girlfriend Charlie less than thrilled). Betty convinces Daniel that he has a sex addiction and needs help.

Unable to decide who their father left in charge of the Meade Empire, Daniel and Alexis settle the matter with a paint ball fight inside MODE offices. Trying to spice her love life, Betty drags Henry into a double date with romantic assertive Gio and her very outgoing sister, Hilda, at a lively reggaeton club. Wilhelmina faces financial problems in new launched Slater Magazine and tries to improve her public image ending in a total media disaster as she ran over Betty White by mistake. Amanda and Nick finally settle their differences.

Daniel struggles to deal with Alex running Meade Publications following their paint-ball showdown, while Betty campaigns for healthier models for this year's fashion week show.

While the whole Meade family is involved in Claire's trial on Fey Sommer's murder, Betty comes upon a perfume that makes her go hyper-mode and drives both Henry and Gio crazy about her. After violently breaking into Gio's deli and ending in a hospital, Betty and her family discover the deadly truth about the mysterious perfume. Amanda has a moral dilemma regarding the past of her late mother and Wilhelmina, with the help of Marc, will try the impossible to make Christina agree to join her in her mischievous plans.