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The story takes place in Japan, apparently where Syaoran and Sakura live. Syaoran and Sakura were in Clamp's very popular anime "Cardcaptors." However in this story, they are different. Sakura is a princess and Syaoran is her childhood friend. They met long ago, and won't admit yet to each other that they could possibly have fallen in love. But a last confession, suddenly takes a sudden turn of events and Sakura ends up losing her memory. To save her, Syaoran bargains their relationship to bring her back, and must journey to many worlds with companions to retrieve her wings (tsubasa) that have taken flight. No matter how painful it might be.

Tsubasa Chronicles has many characters from CLAMP. Including, Syaoran and Sakura. Syaoran lives in the kingdom of Clow, ruled by King Toya. (Fans of Cardcaptor Sakura will know that this is Sakura's brother) After his father died (who actually was Sakura's father in CCS) he still carried on his father's and his dream to uncover the mysterious ruins of Clow. Princess Sakura, of Clow, comes to see Syaoran when he returns from a dig. Later on she tries to tell Syaoran her true feelings, about how she loves him but many things get in the way. That same night Sakura was drawn to the ruins of Clow by a mysterious force. Syaoran was there, examining the ruins, when Sakura came and that's when everything changed. Sakura's feathers, her heart which are her memories, disappeared to many worlds and if they are not found then Princess Sakura will die. Yukito, the head priest, sent Syaoran with Sakura to the space time witch who many fans of CLAMP will know is Yuuko. In other worlds, Kurogane is sent away from Princess Tomoyo's kingdom to Yuuko since he is too violent and she wants him to find more to life. Fai, in another world, just sealed off his enemy and now, leaving Chi in charge of guarding, flees from his world and goes to Yuuko. Each now travels many worlds together, using Mokona to travel to worlds at random. But in order to travel they must all give up something in return. Kurogane gave up his sword, Fai the symbols on his back which is his magic and Syaoran has to give up his relationship with Sakura. Even if Syaoran does success to retrieve Sakura's feathers, she won't be able to remember him. Syaoran still goes through with it since he doesn't want the most important person to him to die. Now, the five of them are traveling to different worlds meeting people they once knew and finding new friends.