The four goddesses of Gamindustri, Neptune, Noire, Blanc, and Vert, have been selected to become beta testers for the hit online game 4 Goddesses Online, with their likenesses used as their game avatars. With the help of the game's virtual guide Bouquet, the four have been chosen to revive the Goddesses of Legend in order to protect the land of Alsguard from the threat of the Demon King Jester. But not just that, but the goddesses must also chase after a mysterious hacker who has been using cheat tactics in order to make the game impossible to play, seeking to destroy the game's popularity.
