For Series 2, each week's five episodes constituted one journey. Episodes Brighton to Crystal Palace (2011) to Dereham to Cromer (2011) formed the first journey, north-eastwards across southern England and East Anglia, from Brighton via London to Cromer. Episodes Ledbury to Shrewsbury (2011) to Llanberis to Holyhead (2011) formed the second journey, westwards across the English Midlands into north Wales, following the route of the historic Irish Mail train, from Ledbury to Holyhead. Episodes Newcastle to Chester-le-Street (2011) to Langley Mill to Melton Mowbray (2011) formed the third journey, southwards across northern England and the Midlands, from Newcastle to Melton Mowbray. Episodes London Bridge to Chatham (2011) to Hythe to Hastings (2011) formed the fourth journey, southwards across southern England to the south coast, from London to Hastings. Episodes Ayr to Paisley (2011) to Lochailort to Skye (2011) formed the fifth journey across Scotland from Ayr to the Isle of Skye.
The opening title sequence for this episode was "Great British Railway Journeys Goes to Ireland".
For Series 1, each week's five episodes constituted one journey. Episodes Liverpool to Eccles (2010) to Filey to Scarborough (2010) formed the first journey, eastwards across northern England, from Liverpool to Scarborough. Episodes Preston to Morecambe (2010) to Edinburgh to Kirkcaldy (2010) formed the second journey, northwards up the west of northern England into Scotland, from Preston to Kirkcaldy. Episodes Swindon to Bristol (2010) to Truro to Penzance (2010) formed the third journey, westwards across southern England, from Swindon to Penzance. Episodes Buxton to Matlock (2010) to St. Pancras to Westminster (2010) formed the fourth journey, southwards across central England, from Buxton to London.